2025 Mary E. Brunkow — for their discoveries concerning peripheral immune tolerance
2025 Fred Ramsdell United States United Statesfor their discoveries concerning peripheral immune tolerance
2025 Shimon Sakaguchi Japan Japanfor their discoveries concerning peripheral immune tolerance
2024 Victor Ambros United States United Statesfor the discovery of microRNA and its role in post-transcriptional gene regulation
2024 Gary Ruvkun United States United Statesfor the discovery of microRNA and its role in post-transcriptional gene regulation
2023 Katalin Karikó United States United Statesfor their discoveries concerning nucleoside base modifications that enabled the development of effective mRNA vaccines against COVID-19
2023 Drew Weissman United States United Statesfor their discoveries concerning nucleoside base modifications that enabled the development of effective mRNA vaccines against COVID-19
2022 Svante Pääbo Sweden Swedenfor his discoveries concerning the genomes of extinct hominins and human evolution
2021 David Julius United States United Statesfor their discoveries of receptors for temperature and touch
2021 Ardem Patapoutian United States United Statesfor their discoveries of receptors for temperature and touch
2020 Harvey J. Alter United States United Statesfor the discovery of Hepatitis C virus
2020 Michael Houghton United Kingdom United Kingdomfor the discovery of Hepatitis C virus
2020 Charles M. Rice United States United Statesfor the discovery of Hepatitis C virus
2019 William G. Kaelin Jr United States United Statesfor their discoveries of how cells sense and adapt to oxygen availability
2019 Sir Peter J. Ratcliffe United Kingdom United Kingdomfor their discoveries of how cells sense and adapt to oxygen availability
2019 Gregg L. Semenza United States United Statesfor their discoveries of how cells sense and adapt to oxygen availability
2018 James P. Allison United States United Statesfor their discovery of cancer therapy by inhibition of negative immune regulation
2018 Tasuku Honjo Japan Japanfor their discovery of cancer therapy by inhibition of negative immune regulation
2017 Jeffrey C. Hall United States United Statesfor their discoveries of molecular mechanisms controlling the circadian rhythm
2017 Michael Rosbash United States United Statesfor their discoveries of molecular mechanisms controlling the circadian rhythm
2017 Michael W. Young United States United Statesfor their discoveries of molecular mechanisms controlling the circadian rhythm
2016 Yoshinori Ohsumi Japan Japanfor his discoveries of mechanisms for autophagy
2015 William C. Campbell United States United Statesfor their discoveries concerning a novel therapy against infections caused by roundworm parasites
2015 Satoshi Ōmura Japan Japanfor their discoveries concerning a novel therapy against infections caused by roundworm parasites
2015 Tu Youyou China Chinafor her discoveries concerning a novel therapy against Malaria
2014 John O'Keefe United Kingdom United Kingdomfor their discoveries of cells that constitute a positioning system in the brain
2014 May-Britt Moser Norway Norwayfor their discoveries of cells that constitute a positioning system in the brain
2014 Edvard I. Moser Norway Norwayfor their discoveries of cells that constitute a positioning system in the brain
2013 James E. Rothman United States United Statesfor their discoveries of machinery regulating vesicle traffic, a major transport system in our cells
2013 Randy W. Schekman United States United Statesfor their discoveries of machinery regulating vesicle traffic, a major transport system in our cells
2013 Thomas C. Südhof United States United Statesfor their discoveries of machinery regulating vesicle traffic, a major transport system in our cells
2012 Sir John B. Gurdon United Kingdom United Kingdomfor the discovery that mature cells can be reprogrammed to become pluripotent
2012 Shinya Yamanaka Japan Japanfor the discovery that mature cells can be reprogrammed to become pluripotent
2011 Bruce A. Beutler United States United Statesfor their discoveries concerning the activation of innate immunity
2011 Jules A. Hoffmann France Francefor their discoveries concerning the activation of innate immunity
2011 Ralph Steinman United States United Statesfor his discovery of the dendritic cell and its role in adaptive immunity
2010 Robert G. Edwards United Kingdom United Kingdomfor the development of in vitro fertilization
2009 Elizabeth Blackburn United States United Statesfor the discovery of how chromosomes are protected by telomeres and the enzyme telomerase
2009 Carol W. Greider United States United Statesfor the discovery of how chromosomes are protected by telomeres and the enzyme telomerase
2009 Jack Szostak United States United Statesfor the discovery of how chromosomes are protected by telomeres and the enzyme telomerase
2008 Harald zur Hausen Germany Germanyfor his discovery of human papilloma viruses causing cervical cancer
2008 Françoise Barré-Sinoussi France Francefor their discovery of human immunodeficiency virus
2008 Luc Montagnier France Francefor their discovery of human immunodeficiency virus
2007 Mario R. Capecchi United States United Statesfor their discoveries of principles for introducing specific gene modifications in mice by the use of embryonic stem cells
2007 Sir Martin J. Evans United Kingdom United Kingdomfor their discoveries of principles for introducing specific gene modifications in mice by the use of embryonic stem cells
2007 Oliver Smithies United States United Statesfor their discoveries of principles for introducing specific gene modifications in mice by the use of embryonic stem cells
2006 Andrew Z. Fire United States United Statesfor their discovery of RNA interference - gene silencing by double-stranded RNA
2006 Craig C. Mello United States United Statesfor their discovery of RNA interference - gene silencing by double-stranded RNA
2005 Barry Marshall Australia Australiafor their discovery of the bacterium <i>Helicobacter pylori</i> and its role in gastritis and peptic ulcer disease
2005 Robin Warren Australia Australiafor their discovery of the bacterium <i>Helicobacter pylori</i> and its role in gastritis and peptic ulcer disease
2004 Richard Axel United States United Statesfor their discoveries of odorant receptors and the organization of the olfactory system
2004 Linda B. Buck United States United Statesfor their discoveries of odorant receptors and the organization of the olfactory system
2003 Paul C. Lauterbur United States United Statesfor their discoveries concerning magnetic resonance imaging
2003 Sir Peter Mansfield United Kingdom United Kingdomfor their discoveries concerning magnetic resonance imaging
2002 Sydney Brenner United Kingdom United Kingdomfor their discoveries concerning genetic regulation of organ development and programmed cell death
2002 H. Robert Horvitz United States United Statesfor their discoveries concerning genetic regulation of organ development and programmed cell death
2002 John E. Sulston United Kingdom United Kingdomfor their discoveries concerning genetic regulation of organ development and programmed cell death
2001 Leland Hartwell United States United Statesfor their discoveries of key regulators of the cell cycle
2001 Tim Hunt United Kingdom United Kingdomfor their discoveries of key regulators of the cell cycle
2001 Sir Paul Nurse United Kingdom United Kingdomfor their discoveries of key regulators of the cell cycle
2000 Arvid Carlsson Sweden Swedenfor their discoveries concerning signal transduction in the nervous system
2000 Paul Greengard United States United Statesfor their discoveries concerning signal transduction in the nervous system
2000 Eric Kandel United States United Statesfor their discoveries concerning signal transduction in the nervous system
1999 Günter Blobel Germany Germanyfor the discovery that proteins have intrinsic signals that govern their transport and localization in the cell
1998 Robert F. Furchgott United States United Statesfor their discoveries concerning nitric oxide as a signalling molecule in the cardiovascular system
1998 Louis J. Ignarro United States United Statesfor their discoveries concerning nitric oxide as a signalling molecule in the cardiovascular system
1998 Ferid Murad United States United Statesfor their discoveries concerning nitric oxide as a signalling molecule in the cardiovascular system
1997 Stanley B. Prusiner United States United Statesfor his discovery of Prions - a new biological principle of infection
1996 Peter C. Doherty Australia Australiafor their discoveries concerning the specificity of the cell mediated immune defence
1996 Rolf M. Zinkernagel Switzerland Switzerlandfor their discoveries concerning the specificity of the cell mediated immune defence
1995 Edward B. Lewis United States United Statesfor their discoveries concerning the genetic control of early embryonic development
1995 Christiane Nüsslein-Volhard Germany Germanyfor their discoveries concerning the genetic control of early embryonic development
1995 Eric F. Wieschaus United States United Statesfor their discoveries concerning the genetic control of early embryonic development
1994 Alfred G. Gilman United States United Statesfor their discovery of G-proteins and the role of these proteins in signal transduction in cells
1994 Martin Rodbell United States United Statesfor their discovery of G-proteins and the role of these proteins in signal transduction in cells
1993 Richard J. Roberts United Kingdom United Kingdomfor their discoveries of split genes
1993 Phillip A. Sharp United States United Statesfor their discoveries of split genes
1992 Edmond H. Fischer United States United Statesfor their discoveries concerning reversible protein phosphorylation as a biological regulatory mechanism
1992 Edwin G. Krebs United States United Statesfor their discoveries concerning reversible protein phosphorylation as a biological regulatory mechanism
1991 Erwin Neher Germany Germanyfor their discoveries concerning the function of single ion channels in cells
1991 Bert Sakmann Germany Germanyfor their discoveries concerning the function of single ion channels in cells
1990 Joseph E. Murray United States United Statesfor their discoveries concerning organ and cell transplantation in the treatment of human disease
1990 E. Donnall Thomas United States United Statesfor their discoveries concerning organ and cell transplantation in the treatment of human disease
1989 J. Michael Bishop United States United Statesfor their discovery of the cellular origin of retroviral oncogenes
1989 Harold E. Varmus United States United Statesfor their discovery of the cellular origin of retroviral oncogenes
1988 Sir James W. Black United Kingdom United Kingdomfor their discoveries of important principles for drug treatment
1988 Gertrude B. Elion United States United Statesfor their discoveries of important principles for drug treatment
1988 George H. Hitchings United States United Statesfor their discoveries of important principles for drug treatment
1987 Susumu Tonegawa Japan Japanfor his discovery of the genetic principle for generation of antibody diversity
1986 Stanley Cohen United States United Statesfor their discoveries of growth factors
1986 Rita Levi-Montalcini United States United Statesfor their discoveries of growth factors
1985 Michael S. Brown United States United Statesfor their discoveries concerning the regulation of cholesterol metabolism
1985 Joseph L. Goldstein United States United Statesfor their discoveries concerning the regulation of cholesterol metabolism
1984 Niels Kaj Jerne Denmark Denmarkfor theories concerning the specificity in development and control of the immune system and the discovery of the principle for production of monoclonal antibodies
1984 Georges J.F. Köhler Germany Germanyfor theories concerning the specificity in development and control of the immune system and the discovery of the principle for production of monoclonal antibodies
1984 César Milstein United Kingdom United Kingdomfor theories concerning the specificity in development and control of the immune system and the discovery of the principle for production of monoclonal antibodies
1983 Barbara McClintock United States United Statesfor her discovery of mobile genetic elements
1982 Sune K. Bergström Sweden Swedenfor their discoveries concerning prostaglandins and related biologically active substances
1982 Bengt I. Samuelsson Sweden Swedenfor their discoveries concerning prostaglandins and related biologically active substances
1982 John R. Vane United Kingdom United Kingdomfor their discoveries concerning prostaglandins and related biologically active substances
1981 Roger W. Sperry United States United Statesfor his discoveries concerning the functional specialization of the cerebral hemispheres
1981 David H. Hubel United States United Statesfor their discoveries concerning information processing in the visual system
1981 Torsten N. Wiesel United States United Statesfor their discoveries concerning information processing in the visual system
1980 Baruj Benacerraf United States United Statesfor their discoveries concerning genetically determined structures on the cell surface that regulate immunological reactions
1980 Jean Dausset France Francefor their discoveries concerning genetically determined structures on the cell surface that regulate immunological reactions
1980 George D. Snell United States United Statesfor their discoveries concerning genetically determined structures on the cell surface that regulate immunological reactions
1979 Allan McLeod Cormack United States United Statesfor the development of computer assisted tomography
1979 Godfrey N. Hounsfield United Kingdom United Kingdomfor the development of computer assisted tomography
1978 Werner Arber Switzerland Switzerlandfor the discovery of restriction enzymes and their application to problems of molecular genetics
1978 Daniel Nathans United States United Statesfor the discovery of restriction enzymes and their application to problems of molecular genetics
1978 Hamilton O. Smith United States United Statesfor the discovery of restriction enzymes and their application to problems of molecular genetics
1977 Roger Guillemin United States United Statesfor their discoveries concerning the peptide hormone production of the brain
1977 Andrew V. Schally Poland Polandfor their discoveries concerning the peptide hormone production of the brain
1977 Rosalyn Yalow United States United Statesfor the development of radioimmunoassays of peptide hormones
1976 Baruch S. Blumberg United States United Statesfor their discoveries concerning new mechanisms for the origin and dissemination of infectious diseases
1976 D. Carleton Gajdusek United States United Statesfor their discoveries concerning new mechanisms for the origin and dissemination of infectious diseases
1975 David Baltimore United States United Statesfor their discoveries concerning the interaction between tumour viruses and the genetic material of the cell
1975 Renato Dulbecco United States United Statesfor their discoveries concerning the interaction between tumour viruses and the genetic material of the cell
1975 Howard M. Temin United States United Statesfor their discoveries concerning the interaction between tumour viruses and the genetic material of the cell
1974 Albert Claude United States United Statesfor their discoveries concerning the structural and functional organization of the cell
1974 Christian de Duve Belgium Belgiumfor their discoveries concerning the structural and functional organization of the cell
1974 George Emil Palade United States United Statesfor their discoveries concerning the structural and functional organization of the cell
1973 Karl von Frisch Austria Austriafor their discoveries concerning organization and elicitation of individual and social behaviour patterns
1973 Konrad Lorenz Austria Austriafor their discoveries concerning organization and elicitation of individual and social behaviour patterns
1973 Niko Tinbergen United Kingdom United Kingdomfor their discoveries concerning organization and elicitation of individual and social behaviour patterns
1972 Gerald M. Edelman United States United Statesfor their discoveries concerning the chemical structure of antibodies
1972 Rodney R. Porter United Kingdom United Kingdomfor their discoveries concerning the chemical structure of antibodies
1971 Earl W. Sutherland, Jr. United States United Statesfor his discoveries concerning the mechanisms of the action of hormones
1970 Sir Bernard Katz United Kingdom United Kingdomfor their discoveries concerning the humoral transmitters in the nerve terminals and the mechanism for their storage, release and inactivation
1970 Ulf von Euler Sweden Swedenfor their discoveries concerning the humoral transmitters in the nerve terminals and the mechanism for their storage, release and inactivation
1970 Julius Axelrod Poland Polandfor their discoveries concerning the humoral transmitters in the nerve terminals and the mechanism for their storage, release and inactivation
1969 Max Delbrück United States United Statesfor their discoveries concerning the replication mechanism and the genetic structure of viruses
1969 Alfred D. Hershey United States United Statesfor their discoveries concerning the replication mechanism and the genetic structure of viruses
1969 Salvador E. Luria United States United Statesfor their discoveries concerning the replication mechanism and the genetic structure of viruses
1968 Robert W. Holley United States United Statesfor their interpretation of the genetic code and its function in protein synthesis
1968 Har Gobind Khorana India Indiafor their interpretation of the genetic code and its function in protein synthesis
1968 Marshall W. Nirenberg United States United Statesfor their interpretation of the genetic code and its function in protein synthesis
1967 Ragnar Granit Sweden Swedenfor their discoveries concerning the primary physiological and chemical visual processes in the eye
1967 Keffer Hartline United States United Statesfor their discoveries concerning the primary physiological and chemical visual processes in the eye
1967 George Wald United States United Statesfor their discoveries concerning the primary physiological and chemical visual processes in the eye
1966 Peyton Rous United States United Statesfor his discovery of tumour-inducing viruses
1966 Charles B. Huggins United States United Statesfor his discoveries concerning hormonal treatment of prostatic cancer
1965 François Jacob France Francefor their discoveries concerning genetic control of enzyme and virus synthesis
1965 André Lwoff France Francefor their discoveries concerning genetic control of enzyme and virus synthesis
1965 Jacques Monod France Francefor their discoveries concerning genetic control of enzyme and virus synthesis
1964 Konrad Bloch Germany Germanyfor their discoveries concerning the mechanism and regulation of the cholesterol and fatty acid metabolism
1964 Feodor Lynen Germany Germanyfor their discoveries concerning the mechanism and regulation of the cholesterol and fatty acid metabolism
1963 John Eccles Australia Australiafor their discoveries concerning the ionic mechanisms involved in excitation and inhibition in the peripheral and central portions of the nerve cell membrane
1963 Alan Hodgkin United Kingdom United Kingdomfor their discoveries concerning the ionic mechanisms involved in excitation and inhibition in the peripheral and central portions of the nerve cell membrane
1963 Andrew Huxley United Kingdom United Kingdomfor their discoveries concerning the ionic mechanisms involved in excitation and inhibition in the peripheral and central portions of the nerve cell membrane
1962 Francis Crick United Kingdom United Kingdomfor their discoveries concerning the molecular structure of nucleic acids and its significance for information transfer in living material
1962 James Watson United States United Statesfor their discoveries concerning the molecular structure of nucleic acids and its significance for information transfer in living material
1962 Maurice Wilkins United Kingdom United Kingdomfor their discoveries concerning the molecular structure of nucleic acids and its significance for information transfer in living material
1961 Georg von Békésy United States United Statesfor his discoveries of the physical mechanism of stimulation within the cochlea
1960 Frank Macfarlane Burnet Australia Australiafor discovery of acquired immunological tolerance
1960 Peter Medawar United Kingdom United Kingdomfor discovery of acquired immunological tolerance
1959 Severo Ochoa United States United Statesfor their discovery of the mechanisms in the biological synthesis of ribonucleic acid and deoxyribonucleic acid
1959 Arthur Kornberg United States United Statesfor their discovery of the mechanisms in the biological synthesis of ribonucleic acid and deoxyribonucleic acid
1958 George Beadle United States United Statesfor their discovery that genes act by regulating definite chemical events
1958 Edward Tatum United States United Statesfor their discovery that genes act by regulating definite chemical events
1958 Joshua Lederberg United States United Statesfor his discoveries concerning genetic recombination and the organization of the genetic material of bacteria
1957 Daniel Bovet Italy Italyfor his discoveries relating to synthetic compounds that inhibit the action of certain body substances, and especially their action on the vascular system and the skeletal muscles
1956 André F. Cournand United States United Statesfor their discoveries concerning heart catheterization and pathological changes in the circulatory system
1956 Werner Forssmann Germany Germanyfor their discoveries concerning heart catheterization and pathological changes in the circulatory system
1956 Dickinson W. Richards United States United Statesfor their discoveries concerning heart catheterization and pathological changes in the circulatory system
1955 Hugo Theorell Sweden Swedenfor his discoveries concerning the nature and mode of action of oxidation enzymes
1954 John F. Enders United States United Statesfor their discovery of the ability of poliomyelitis viruses to grow in cultures of various types of tissue
1954 Thomas H. Weller United States United Statesfor their discovery of the ability of poliomyelitis viruses to grow in cultures of various types of tissue
1954 Frederick C. Robbins United States United Statesfor their discovery of the ability of poliomyelitis viruses to grow in cultures of various types of tissue
1953 Hans Krebs United Kingdom United Kingdomfor his discovery of the citric acid cycle
1953 Fritz Lipmann United States United Statesfor his discovery of co-enzyme A and its importance for intermediary metabolism
1952 Selman A. Waksman Russia Russiafor his discovery of streptomycin, the first antibiotic effective against tuberculosis
1951 Max Theiler United States United Statesfor his discoveries concerning yellow fever and how to combat it
1950 Edward C. Kendall United States United Statesfor their discoveries relating to the hormones of the adrenal cortex, their structure and biological effects
1950 Tadeusz Reichstein Switzerland Switzerlandfor their discoveries relating to the hormones of the adrenal cortex, their structure and biological effects
1950 Philip S. Hench United States United Statesfor their discoveries relating to the hormones of the adrenal cortex, their structure and biological effects
1949 Walter Hess Switzerland Switzerlandfor his discovery of the functional organization of the interbrain as a coordinator of the activities of the internal organs
1949 António Egas Moniz Portugal Portugalfor his discovery of the therapeutic value of leucotomy in certain psychoses
1948 Paul Müller Switzerland Switzerlandfor his discovery of the high efficiency of DDT as a contact poison against several arthropods
1947 Carl Cori Austria Austriafor their discovery of the course of the catalytic conversion of glycogen
1947 Gerty Cori Austria Austriafor their discovery of the course of the catalytic conversion of glycogen
1947 Bernardo Houssay Argentina Argentinafor his discovery of the part played by the hormone of the anterior pituitary lobe in the metabolism of sugar
1946 Hermann Joseph Muller United States United Statesfor the discovery of the production of mutations by means of X-ray irradiation
1945 Alexander Fleming United Kingdom United Kingdomfor the discovery of penicillin and its curative effect in various infectious diseases
1945 Ernst B. Chain United Kingdom United Kingdomfor the discovery of penicillin and its curative effect in various infectious diseases
1945 Howard Florey Australia Australiafor the discovery of penicillin and its curative effect in various infectious diseases
1944 Joseph Erlanger United States United Statesfor their discoveries relating to the highly differentiated functions of single nerve fibres
1944 Herbert S. Gasser United States United Statesfor their discoveries relating to the highly differentiated functions of single nerve fibres
1943 Henrik Dam Denmark Denmarkfor his discovery of vitamin K
1943 Edward A. Doisy United States United Statesfor his discovery of the chemical nature of vitamin K
1939 Gerhard Domagk Germany Germanyfor the discovery of the antibacterial effects of prontosil
1938 Corneille Heymans Belgium Belgiumfor the discovery of the role played by the sinus and aortic mechanisms in the regulation of respiration
1937 Albert Szent-Györgyi United States United Statesfor his discoveries in connection with the biological combustion processes, with special reference to vitamin C and the catalysis of fumaric acid
1936 Sir Henry Dale United Kingdom United Kingdomfor their discoveries relating to chemical transmission of nerve impulses
1936 Otto Loewi United States United Statesfor their discoveries relating to chemical transmission of nerve impulses
1935 Hans Spemann Germany Germanyfor his discovery of the organizer effect in embryonic development
1934 George H. Whipple United States United Statesfor their discoveries concerning liver therapy in cases of anaemia
1934 George R. Minot United States United Statesfor their discoveries concerning liver therapy in cases of anaemia
1934 William P. Murphy United States United Statesfor their discoveries concerning liver therapy in cases of anaemia
1933 Thomas H. Morgan United States United Statesfor his discoveries concerning the role played by the chromosome in heredity
1932 Sir Charles Sherrington United States United Statesfor their discoveries regarding the functions of neurons
1932 Edgar Adrian United Kingdom United Kingdomfor their discoveries regarding the functions of neurons
1931 Otto Warburg Germany Germanyfor his discovery of the nature and mode of action of the respiratory enzyme
1930 Karl Landsteiner United States United Statesfor his discovery of human blood groups
1929 Christiaan Eijkman Netherlands Netherlandsfor his discovery of the antineuritic vitamin
1929 Sir Frederick Hopkins United Kingdom United Kingdomfor his discovery of the growth-stimulating vitamins
1928 Charles Nicolle France Francefor his work on typhus
1927 Julius Wagner-Jauregg Austria Austriafor his discovery of the therapeutic value of malaria inoculation in the treatment of dementia paralytica
1926 Johannes Fibiger Denmark Denmarkfor his discovery of the Spiroptera carcinoma
1924 Willem Einthoven Netherlands Netherlandsfor his discovery of the mechanism of the electrocardiogram
1923 Frederick Banting Canada Canadafor the discovery of insulin
1923 John Macleod United Kingdom United Kingdomfor the discovery of insulin
1922 Otto Meyerhof United States United Statesfor his discovery of the fixed relationship between the consumption of oxygen and the metabolism of lactic acid in the muscle
1922 Archibald V. Hill United Kingdom United Kingdomfor his discovery relating to the production of heat in the muscle
1920 August Krogh Denmark Denmarkfor his discovery of the capillary motor regulating mechanism
1919 Jules Bordet Belgium Belgiumfor his discoveries relating to immunity
1914 Robert Bárány Sweden Swedenfor his work on the physiology and pathology of the vestibular apparatus
1913 Charles Richet France Francein recognition of his work on anaphylaxis
1912 Alexis Carrel France Francein recognition of his work on vascular suture and the transplantation of blood vessels and organs
1911 Allvar Gullstrand Sweden Swedenfor his work on the dioptrics of the eye
1910 Albrecht Kossel Germany Germanyin recognition of the contributions to our knowledge of cell chemistry made through his work on proteins, including the nucleic substances
1909 Emil Theodor Kocher Switzerland Switzerlandfor his work on the physiology, pathology and surgery of the thyroid gland
1908 Ilya Mechnikov Russia Russiain recognition of their work on immunity
1908 Paul Ehrlich Germany Germanyin recognition of their work on immunity
1907 Charles Louis Alphonse Laveran France Francein recognition of his work on the role played by protozoa in causing diseases
1906 Camillo Golgi Italy Italyin recognition of their work on the structure of the nervous system
1906 Santiago Ramón y Cajal Spain Spainin recognition of their work on the structure of the nervous system
1905 Robert Koch Germany Germanyfor his investigations and discoveries in relation to tuberculosis
1904 Ivan Pavlov Russia Russiain recognition of his work on the physiology of digestion, through which knowledge on vital aspects of the subject has been transformed and enlarged
1903 Niels Ryberg Finsen Denmark Denmarkin recognition of his contribution to the treatment of diseases, especially lupus vulgaris, with concentrated light radiation, whereby he has opened a new avenue for medical science
1902 Ronald Ross United Kingdom United Kingdomfor his work on malaria, by which he has shown how it enters the organism and thereby has laid the foundation for successful research on this disease and methods of combating it
1901 Emil von Behring Germany Germanyfor his work on serum therapy, especially its application against diphtheria, by which he has opened a new road in the domain of medical science and thereby placed in the hands of the physician a victorious weapon against illness and deaths